All of the national newspaper circulaton figures for April, complete with comparisons, can be viewed by clicking on the interactive images below.
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ZenithOptimedia, the Publicis Groupe media network, has downgraded its global ad forecast as a result of disruption to trade and investment caused by tension over Ukraine.
Mail Online continues to lead the websites for the UK's national newspapers, with more than 11.3 million daily unique browsers in August, while the biggest risers year on year are sites for the Metro (175%) and the Mirror (99%).
Trinity Mirror has poached Piers North from Yahoo to take the new role of digital strategy director at the national and regional newspaper publisher.