All of the national newspaper circulaton figures for April, complete with comparisons, can be viewed by clicking on the interactive images below.
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Mail Online continues to lead the websites for the UK's national newspapers, with more than 11.3 million daily unique browsers in August, while the biggest risers year on year are sites for the Metro (175%) and the Mirror (99%).
Trinity Mirror has poached Piers North from Yahoo to take the new role of digital strategy director at the national and regional newspaper publisher.
Starcom MediaVest Group has appointed Sue Frogley to the newly created role of global commercial director.